Or about how something works: Comprehension usually used with of. Understanding implies abilities and dispositions with respect to an object.
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Knowledge about a subject, situation, etc. The meaning of understanding is a mental grasp : How to use understanding in a sentence.
Understanding is a relation between the knower and an object of understanding.
He noted bond's apparent lack of. Both comprehension and understanding can be used to talk about someone's ability to understand something. Your understanding might be that your mother will always drive you to school if you miss the bus. Understanding a concept means you get it.