Mental process of a person who comprehends; He noted bond's apparent lack of. How to use understanding in a sentence.
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Or about how something works: The meaning of understanding is a mental grasp : Both comprehension and understanding can be used to talk about someone's ability to understand something.
Comprehension usually used with of.
Understanding implies abilities and dispositions with respect to an object. Understanding is a relation between the knower and an object of understanding. Knowledge about a subject, situation, etc.