Your understanding might be that your mother will always drive you to school if you miss the bus. The meaning of understanding is a mental grasp : Understanding a concept means you get it.
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How to use understanding in a sentence. Or about how something works: Knowledge about a subject, situation, etc.
Understanding is a relation between the knower and an object of understanding.
Comprehension usually used with of. Understanding implies abilities and dispositions with respect to an object. He noted bond's apparent lack of. Both comprehension and understanding can be used to talk about someone's ability to understand something.