Or about how something works: He noted bond's apparent lack of. Understanding implies abilities and dispositions with respect to an object.
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Understanding a concept means you get it. Knowledge about a subject, situation, etc. How to use understanding in a sentence.
Understanding is a relation between the knower and an object of understanding.
Comprehension usually used with of. Your understanding might be that your mother will always drive you to school if you miss the bus. The meaning of understanding is a mental grasp : Both comprehension and understanding can be used to talk about someone's ability to understand something.